Y-DNA Haplogroup I and its Subclades - 2016

The entire work is identified by the Version Number and date given on the Main Page.   Directions for citing the document are given at the bottom of the Main Page. Version History     Last revision date for this specific page: 11 July 2016

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes and ISOGG does its best to keep the tree updated with the latest developments in the field. The viewer may observe other versions of the tree on the Web. Email Ray Banks if the differences need clarification or if you find any broken links on this page.

LINKS:  Main Page   Y-DNA Tree Trunk   SNP Index   Papers/Presentations Cited   Glossary   Listing Criteria

SNP SYMBOLS:  Not on 2015 tree  Confirmed within subclade  Investigational items

^ Indicates a next-generation sequencing entry which does not yet meet quality guidelines for minimum number of reads.

^^ Indicates an entry which does not meet quality guidelines but may be helpful.

~ Indicates only an approximate location on the tree.

The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing one in multiple labs, and/or being found in the area of the chromosome used in recent research studies.

SNPs listed below in italics (colored black or red) are quality variants from next-generation sequencing reports consistently showing as representing that subgroup.

Contact People for Haplogroup I:
Overall I Contact: Phil Goff
I1 and Subclades Contact: Steve Trangsrud
I2a1 P37.2 and Subclades Contact: Zdenko Markovic
I2a2 P217 and Subclades Contact: Aaron Salles Torres
I2b L415 and I2c L596 Subclades Contact: Bob May

Link to Experimental Composite Y-DNA Haplogroup I Tree by Ray Banks.

I   M170/PF3715, CTS48/PF3569, CTS70/PF3570, CTS88/PF3571, CTS646/PF3629, CTS674/FI1/PF3630, CTS772/PF3631, CTS1006, CTS1301/PF3635, CTS1555/PF3696, CTS1800/PF3699, CTS2193/PF3703, CTS2387/PF3705, CTS2514/PF3706, CTS2536/PF3707, CTS3076/PF3712, CTS3383/PF3716, CTS3384/PF3717, CTS3517/PF3719/V2740, CTS3641/PF3722, CTS4077/PF3725, CTS4088/PF3868, CTS4209/PF3726, CTS4239/PF3727, CTS4272/PF3728, CTS4273/PF3729, CTS4340/PF3730, CTS4637, CTS4664, CTS4745/PF3734, CTS4752/PF3735, CTS4848/FI7/PF3736, CTS4982/PF3737, CTS5016/PF3738, CTS5150/PF3739, CTS5263, CTS5622/PF3743, CTS5650/PF3744, CTS5764/PF3746, CTS5908/PF3747, CTS5946/PF3748, CTS6231/PF3750, CTS6265/PF3871, CTS6271/PF3751, CTS6334/PF3752, CTS6343/PF3753, CTS6344/PF3754, CTS6497/PF3756, CTS6751/PF3757, CTS7026/PF3758, CTS7329, CTS7469, CTS7502/PF3760, CTS7540/PF3761, CTS7593/PF3763, CTS7831/PF3766, CTS8064/PF3768, CTS8257, CTS8300/PF3770, CTS8333/PF3771, CTS8345/PF3772, CTS8420/PF3773, CTS8545/PF3775, CTS8742/PF3776, CTS8876/PF3778, CTS8963, CTS9264/PF3782, CTS9269/PF3783, CTS9484/PF3785, CTS9618/PF3786, CTS9838/PF3788, CTS9860/PF3790, CTS10058, CTS10941/PF3838, CTS11369/PF3840, CTS11441, CTS11540/PF3842, CTS11779/PF3844, CTS11979/PF3878, FI2/YSC0000281, FI3/L756/PF3662/YSC0000284, FI4/PF3668, FGC2411/Y1877, FGC2412/Y1909, FGC2413/Y1869, FGC2414/Y1903, FGC2415/Y1880, FGC2416/S10441/Y1865, FGC2417, FGC2418/Y1841, FGC2419, FGC2420/Y1907, FGC2421/Y1931, FGC2424, FGC2443/Y1834, FGC2447, FGC2453, FGC2456, FGC2458/Y1942, FGC2467, FGC3436, FGC7049, FGC7050, L41/PF3787, L503, L578/PF3653, L751/PF3779/YSC0000291, L755/PF3659/YSC0000283, L758/PF3663/YSC0000285, L772/PF3731/YSC0000263, L844.1/PF3572.1/YSC0000275.1, L846/PF3646/YSC0000280, L847/PF3841/YSC0000298, L1197/PF3718/YSC0000260, M258/PF3721, M11064/PF3634, P19_1, P19_2, P19_3, P19_4, P19_5, P38, P212/PF3580, PF3742/U179, PF3574, PF3575, PF3578, PF3585, PF3586, PF3588, PF3589, PF3590, PF3592, PF3594, PF3596, PF3599, PF3600, PF3601, PF3602, PF3603, PF3604, PF3605, PF3611, PF3612, PF3616, PF3617, PF3618, PF3620, PF3622, PF3625, PF3627.2/V218.2, PF3639, PF3640, PF3641, PF3642, PF3645, PF3649, PF3654, PF3660, PF3661, PF3665, PF3666, PF3670, PF3672, PF3675, PF3677, PF3685, PF3687, PF3689, PF3694, PF3695, PF3742/U179, PF3759/YSC0000267, PF3780, PF3794, PF3795, PF3796, PF3797, PF3799, PF3800, PF3803, PF3804, PF3806, PF3807, PF3809, PF3811, PF3813, PF3814, PF3815, PF3817, PF3819, PF3822, PF3828, PF3829, PF3833, PF3836, PF3837, PF3847/YSC0000300, PF3864, Y1847, YSC0000256.1, YSC0000272, Z16984, Z16985, Z16986/ZS2286, Z16987
• I1   M253, AM00084/CTS2044, CTS10/Z2676, CTS22/Z2677, CTS40/Z2678, CTS136/Z2679, CTS209^/V1092^/Z2680^, CTS565/Z2700, CTS571/Z2701, CTS641/Z2702, CTS733/Z2703, CTS765^/Z2704^, CTS883/Z2705, CTS1220/Z2706, CTS1266/Z2707, CTS1393/Z2708, CTS1739/Z2748, CTS1748/Z2749, CTS1755^/Z2750^, CTS1805/Z2752, CTS2375^/Z2754^, CTS2410/Z2755, CTS2644/Z2757, CTS2732^/Z2758^, CTS2738^/Z2759^, CTS2970/Z2760, CTS3268/Z2762, CTS3474/V2712/Z2764, CTS3506/Z2765, CTS3762/Z2766, CTS3843/Z2767, CTS4037^/V2902^/Z2768^, CTS4130/V2932/Z2769, CTS4189/Z2770, CTS4295/Z2771, CTS4327/Z2772, CTS4342/Z2773, CTS4344/Z2774, CTS4387, CTS4400/Z2775, CTS4508/Z2776, CTS4532^/Z2777^, CTS4612^/Z2778^, CTS4656/Z2779, CTS4675^/Z2780^, CTS4790/Z2781, CTS5167/Z2784, CTS5247/Z2785, CTS5408/Z2786, CTS5513^/Z2787^, CTS5643^/Z2790^, CTS5705/Z2791, CTS5783/S63, CTS5891/Z2793, CTS5993/Z2794, CTS6006/Z2795, CTS6022/Z2796, CTS6076^/Z2797^, CTS6109/Z2798, CTS6140/Z2799, CTS6221^/Z2802^, CTS6395/Z2803, CTS6547/Z2804, CTS6629/Z2805, CTS6765/Z2806, CTS6992, CTS7061/Z2807, CTS7121/Z2808, CTS7267^/Z2809^, CTS7386/Z2810, CTS7477/Z2811, CTS7652/Z2812, CTS7751^/Z2813^, CTS7949^/Z2814^, CTS8170/Z2816, CTS8224/Z2817, CTS8394/Z2818, CTS8652/Z2820, CTS8708/Z2821, CTS8716^/Z2822^, CTS9010/Z2824, CTS9258/Z2825, CTS9288/Z2826, CTS9368^/Z2827^, CTS9405/Z2828, CTS9486/Z2829, CTS9487^/Z2830^, CTS9518/Z2831, CTS9648^/Z2832^, CTS9749/Z2833, CTS9900^, CTS10062/Z2834, CTS10140/Z2837, CTS10338, CTS10613/Z2840, CTS11036/Z2863, CTS11042/S66, CTS11091/Z2865, CTS11126^/Z2866^, CTS11166^/Z2867^, CTS11214/Z2868, CTS11503^/Z2869^, CTS11526/Z2870, CTS11534/Z2871, CTS11552/Z2872, CTS11751/Z2874, CTS11775/Z2875, CTS11783/Z2876, CTS11838/Z2879, CTS11950^/Z1880^, CTS12008/Z2881, F3033.2/M1443.2/V4056.2/Z2838.2, FGC2422/Z2715, FGC2425/Z2714, FGC2427^/Z2713^, FGC2430/Z2699, FGC2431/Z2710, FGC2432/V1339/Z2712, FGC2435/Z2698, FGC2436/Y1879, FGC2439/Z2711, FGC2440/V1427/Y3658, FGC2471/Y1882, FGC2483^/Y1933^, L64, L75, L80, L81, L118, L121/S62, L123, L124/S64, L125/S65, L157.1, L186, L187, L495/Z2788, L509/Z2864, L575/V4059/Z2839, L740^/V3959^/YSC0000296^/Z2836^, L750/YSC0000292/Z2725, L764/YSC0000271/Z2849, L840, L841/YSC0000257/Z2751, L842/YSC0000264/Z2789, L843/YSC0000273/Z2878, L845/S110/YSC0000278, L848/YSC0000299/Z2877, M307.2/P203.2, M450/S109, P30, P40, Page123/Z2763, S107, S108, S111/Y1868, S22865/Z2845, V1423/Z2717, V1771/Z2724, V2264^/Z2728^, Y1886^, YSC0000259/Z2761, YSC0000301/Z2882, Z2718, Z2719, Z2721, Z2722, Z2723, Z2726, Z2727, Z2729, Z2730^, Z2731, Z2733, Z2736, Z2737, Z2738, Z2740, Z2741^, Z2742^, Z2743, Z2744, Z2747, Z2823, Z2841, Z2842, Z2843, Z2844, Z2846, Z2847, Z2848, Z2850, Z2851^, Z2852^, Z2853^, Z2854^, Z2855^, Z2856, Z2857^, Z2861, Z2862, Z2885, Z2887
• I1~ L234, L410, L1274, L1275
• • I1a   DF29/S438
• • • I1a1   CTS6364/Z2336
• • • • I1a1a   M227
• • • • • I1a1a1   M72
• • • • I1a1b   L22/S142
• • • • I1a1b~S336/Z75
• • • • • I1a1b1   P109
• • • • • I1a1b2   L205.1/L939.1/S239.1
• • • • • I1a1b3   Z74
• • • • • • I1a1b3a   L287/S240
• • • • • • • I1a1b3a1   L258/S335
• • • • • • • • I1a1b3a1a   L296
• • • • • • I1a1b3b   L813/S436/Z719
• • • • • I1a1b4   L300/S241
• • • • • I1a1b5~S297/Z718
• • • • • I1a1b6~L242
• • • I1a2    S244/Z58
• • • • I1a2a   S246/Z59
• • • • • I1a2a1   S337/Z60, S439/Z61, Z62
• • • • • • I1a2a1a   S440/Z140, Z141
• • • • • • • I1a2a1a1   S1953/Z2535
• • • • • • • • I1a2a1a1a   S1954/YSC0000261
• • • • • • • • • I1a2a1a1a1   L338/S197
• • • • • • • • • I1a2a1a1a1~ A376
• • • • • • • • I1a2a1a1b   CTS10937/S4698/Z2538
• • • • • • • I1a2a1a2   F2642.1/S2169.1
• • • • • • • I1a2a1a3   A196
• • • • • • • • I1a2a1a3a   Y6900
• • • • • • • • • I1a2a1a3a1   Y7140
• • • • • • • • • • I1a2a1a3a1a   Y7477
• • • • • • • • • • • I1a2a1a3a1a1   YP1081
• • • • • • • • • I1a2a1a3a2   Y6885
• • • • • • I1a2a1b   S247/Z73
• • • • • • • I1a2a1b1   BY266/Y3657
• • • • • • • • I1a2a1b1a   L1302
• • • • • • • • I1a2a1b1a~ BY148, BY211, BY212, BY213, BY214, BY215, BY216, BY217, BY218, BY220, BY222, BY223, BY224, BY225, BY226, BY228, BY230, BY232, BY233, BY234, BY235, BY236, BY240, BY244, BY245, BY269, BY270, BY271, BY272, BY273, BY274, BY275, BY276, BY277, BY278
• • • • • • • • • I1a2a1b1a1   BY147
• • • • • • • • • I1a2a1b1a2   BY126
• • • • • • I1a2a1c   L573
• • • • • • I1a2a1d   L1248
• • • • • • • I1a2a1d1   L803
• • • • • • I1a2a1e~ CTS7362/S4713/Z2539
• • • • • I1a2a2   Z382
• • • • • I1a2a2~ L1450, L1451, L1452, Z2037, Z2038, Z2040, Z2041
• • • • I1a2b   S296/Z138, Z139/S338, PF2805.2/Z2540.2
• • • • • I1a2b1   Z2541
• • • I1a3   S243/Z63
• • • I1a3~ BY12, BY13
• • • • I1a3a   BY151
• • • • • I1a3a1   L849.2
• • • • • • I1a3a1a   S2078
• • • • • • • I1a3a1a1   Y2245.2
• • • • • • • • I1a3a1a1a   L1237
• • • • • • • • • I1a3a1a1a1   FGC9550/Y6634
• • • • • • • • I1a3a1a1b   S10360, S17023, S26266
• • • • • • • • • I1a3a1a1b1   S15301
• • • • • • • • • • I1a3a1a1b1a   Y6228
• • • • • • • • • • • I1a3a1a1b1a1   Y6229
• • • • • • • • • • I1a3a1a1b1b   Y3969
• • • • • • • • I1a3a1a1c   CTS3848, FGC14479
• • • • • • • I1a3a1a2   Y6375, Y6383
• • • • • I1a3a2   BY351, BY352
• • • • • • I1a3a2a   BY332, BY354/ Y7075
• • • • • • • I1a3a2a1   Y7060, Y7059
• • • • • • • • I1a3a2a1a   BY11, S11542.1/V2110.1
• • • • • • • I1a3a2a2   Y13946, PH1538/Y14110
• • • • • • I1a3a2b   CTS10345, CTS2910/S4736
• • • • • • • I1a3a2b1   Y10994
• • • • • • I1a3a2c   CTS4279
• • • • • I1a3a3   BY62
• • I1b   S249/Z131
• • • I1b1   CTS6397
• • I1c   Y18119/Z17925, S2304/Z17937, Z17943
• I2   M438/P215/PF3853/S31, L68/PF3781/S329, PF3664/Z2638
• • I2a   L460/PF3647/S238
• • • I2a1   P37.2/PF4004
• • • • I2a1a   CTS595
• • • • • I2a1a1   M26/PF4056, L158/PF4073/S433, L159.1/S169.1
• • • • • I2a1a1~L673
• • • • • • I2a1a1a   L672/S327
• • • • • • • I2a1a1a1   L160/PF4013/S184
• • • • • • • I2a1a1a1~ Z109, Z112, Z124
• • • • • • • • I2a1a1a1a   PF4088/S5312
• • • • • • • • • I2a1a1a1a1   CTS11338/PF4194
• • • • • • • • • • I2a1a1a1a1a   PF4189
• • • • • • • • • • I2a1a1a1a1b   Z118
• • • • • • • • • • • I2a1a1a1a1b1~ S294/Z106
• • • • • • • • I2a1a1a1b   F1295
• • • • • • • • I2a1a1a1c~ Z119, Z125
• • • • • I2a1a2   S21825
• • • • • • I2a1a2a   L1286
• • • • • • • I2a1a2a1   L1287
• • • • • • • • I2a1a2a1a   L233/S183
• • • • • • • • • I2a1a2a1a1   A417
• • • • • • • I2a1a2a2   L880
• • • • • • I2a1a2b   L1294
• • • • I2a1b   M423, CTS176/S2621, CTS1293/S2632, CTS1802/S2638, CTS5375/S2679, CTS5985/S2687, CTS7218/S2702, CTS8239/S2715, CTS8486/S2722, CTS11030/S2768, L178/S328
• • • • I2a1b~L1224
• • • • • I2a1b1   L161.1/S185
• • • • • • I2a1b1a   L1498
• • • • • • • I2a1b1a1   S2703
• • • • • • I2a1b1b   PF4135
• • • • • • I2a1b1b~ A813, A814
• • • • • I2a1b2   L621/S392
• • • • • • I2a1b2a   CTS10936
• • • • • • • I2a1b2a1   CTS5966, CTS10228, L147.2
• • • • • • • I2a1b2a1~ A493
• • • • • • • • I2a1b2a1a   S17250/YP204
• • • • • • • • • I2a1b2a1a1   Z16971
• • • • • • • • • I2a1b2a1a1~ BY128
• • • • • • • • • • I2a1b2a1a1a~A815
• • • • • • • • • I2a1b2a1a2   Y4882
• • • • • • • • • • I2a1b2a1a2a~ A811
• • • • • • • • • I2a1b2a1a3   A356/Z16983
• • • • • • • • • I2a1b2a1a4~ M359.2/P41.2
• • • • • • • • I2a1b2a1b   Y4460
• • • • • • • • • I2a1b2a1b1   Y3118
• • • • • • • • I2a1b2a1c   Z17855
• • • I2a2   M436/P214/PF3856/S33, L35/PF3862/S150, L37/PF6900/S153, L181, L368, P216/PF3855/S30, P217/PF3854/S23, P218/S32
• • • • I2a2a   M223, L34/PF3857/S151, L36/S152, L59, L622, P219/PF3859/S24, P220/S119, P221/PF3858/S120, P222/PF3861/U250/S118, P223/PF3860/S117, Z77
• • • • I2a2a~ L387.2, L700, L705.1, L706.1
• • • • • I2a2a1   CTS616, CTS9183
• • • • • • I2a2a1a   FGC15073/Y3721
• • • • • • • I2a2a1a1   M284
• • • • • • • • I2a2a1a1a   L1195 • • • • • • • • • I2a2a1a1a1   L126/S165, L137.1/S166.1, L369
• • • • • • • • • • I2a2a1a1a1a   FGC20048/S7753/Y4171
• • • • • • • • • • • I2a2a1a1a1a1   Y8701
• • • • • • • • • • • • I2a2a1a1a1a1a   Y4751
• • • • • • • • • • • • • I2a2a1a1a1a1a1   Y10655
• • • • • • • • • • • • • I2a2a1a1a1a1a2   Y4752
• • • • • • • • • • • • • • I2a2a1a1a1a1a2a   Y4750
• • • • • • • • • • • • • • • I2a2a1a1a1a1a2a1   A824/Y11197
• • • • • • • • • • • • • • • I2a2a1a1a1a1a2a2   Y5673
• • • • • • • • • • • • • • • • I2a2a1a1a1a1a2a2a   Y11246, S15280.2
• • • • • • • • • • • • • • • I2a2a1a1a1a1a2a3   Y8599
• • • • • • • • • • • • I2a2a1a1a1a1b   Y7190
• • • • • • • • • I2a2a1a1a2   L1193
• • • • • • • • • • I2a2a1a1a2a   CTS4922, Y3684, Y3713
• • • • • • • I2a2a1a2   Z2057, Z2083, Z2093
• • • • • • • • I2a2a1a2a   L1229
• • • • • • • • • I2a2a1a2a1   Y3681, Z2069
• • • • • • • • • • I2a2a1a2a1a   Z2058
• • • • • • • • • • • I2a2a1a2a1a1   Z2054
• • • • • • • • • • • I2a2a1a2a1a1~ BY18
• • • • • • • • • • • • I2a2a1a2a1a1a   BY524/Y9443
• • • • • • • • • • • • • I2a2a1a2a1a1a1   L812/S391
• • • • • • • • • • • • • • I2a2a1a2a1a1a1a   Y5308, Y5318
• • • • • • • • • • • • • • I2a2a1a2a1a1a1b   Y13008
• • • • • • • • • • • • • I2a2a1a2a1a1a2   Y10648
• • • • • • • • • • • • I2a2a1a2a1a1b   Y7244
• • • • • • • • • • • • • I2a2a1a2a1a1b1   P53.3
• • • • • • • • • • • • • I2a2a1a2a1a1b2   Y7243
• • • • • • • • • • • • I2a2a1a2a1a1c   FGC15106/Y4334, Y4332
• • • • • • • • • • • • • I2a2a1a2a1a1c1   Y4760
• • • • • • • • • • • • I2a2a1a2a1a1d   BY138
• • • • • • • • • • • • • I2a2a1a2a1a1d1   Y13321
• • • • • • • • • I2a2a1a2a2   S18331, SK1247/Y4915
• • • • • • • • • • I2a2a1a2a2a   L1230
• • • • • • • • I2a2a1a2b   Y7240
• • • • • • I2a2a1b   CTS10057, CTS10100
• • • • • • • I2a2a1b1   L701, L702
• • • • • • • • I2a2a1b1a   P78
• • • • • • • • • I2a2a1b1a1   S25733
• • • • • • • • • • I2a2a1b1a1a   A427
• • • • • • • • • • • I2a2a1b1a1a1   S23612, Y4884
• • • • • • • • • • • • I2a2a1b1a1a1a   Y5360, Y5359
• • • • • • • • • • • • • I2a2a1b1a1a1a1   Y5369
• • • • • • • • • • • • I2a2a1b1a1a1b   S10702, Y6396
• • • • • • • • • • • • • I2a2a1b1a1a1b1   Y11261
• • • • • • • • • I2a2a1b1a2   Y7219
• • • • • • • • • • I2a2a1b1a2a   Y7214
• • • • • • • • • • I2a2a1b1a2b   Y8945
• • • • • • • • I2a2a1b1b   L699, L703
• • • • • • • • •  I2a2a1b1b1   L704
• • • • • • • • • • I2a2a1b1b1a   Y7641
• • • • • • • • •  I2a2a1b1b2   S12195, S21579, Y5669
• • • • • • • • • • I2a2a1b1b2a   Y6973, BY159/Y6996
• • • • • • • I2a2a1b2   Z161
• • • • • • • • I2a2a1b2a   L801/S390
• • • • • • • • •  I2a2a1b2a1   CTS1977
• • • • • • • • • • I2a2a1b2a1a   Y5282
• • • • • • • • • • • I2a2a1b2a1a1   S8522
• • • • • • • • • • • • I2a2a1b2a1a1a   P95
• • • • • • • • • • • I2a2a1b2a1a2   CTS1858
• • • • • • • • • • • • I2a2a1b2a1a2a   CTS10148
• • • • • • • • • • I2a2a1b2a1b   Y7152, S16196
• • • • • • • • • • I2a2a1b2a1c   BY526/Y8935
• • • • • • • • •  I2a2a1b2a2   CTS6433
• • • • • • • • • • I2a2a1b2a2a   S2364
• • • • • • • • • • • I2a2a1b2a2a1   S2361, Z167
• • • • • • • • • • • • I2a2a1b2a2a1a   Z78
• • • • • • • • • • • • • I2a2a1b2a2a1a1   CTS8584
• • • • • • • • • • • • • • I2a2a1b2a2a1a1a   Z185
• • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1   Z180
• • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a   L1198
• • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a1   S20905
• • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a1a   Z190
• • • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a1a1   S434/Z79
• • • • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a1a1a   Y5729, Y5730
• • • • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a1a1b   PH3769/Y7682
• • • • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a1a1c   Y8712
• • • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a1a2   Y7280
• • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a2   P195.2
• • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a3   Y6060
• • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a3a   Y5748
• • • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a3a1   Y7272
• • • • • • • • • • • • • • • • • • I2a2a1b2a2a1a1a1a3b   Y7273
• • • • • • • • • • • • I2a2a1b2a2a1b   Y9161
• • • • • • • • • • • I2a2a1b2a2a2   ZS20/Y4925, S8112
• • • • • • • • • • • • I2a2a1b2a2a2a   CTS661/L1272
• • • • • • • • • • • • • I2a2a1b2a2a2a1   Y5717
• • • • • • • • • • • • I2a2a1b2a2a2b   Y4926
• • • • • • • • • • • • • I2a2a1b2a2a2b1   S8104, Y6883
• • • • • • • • • • • I2a2a1b2a2a3   Y4955
• • • • • • • • • • • • I2a2a1b2a2a3a   Y5695
• • • • • • • • • • • • • I2a2a1b2a2a3a1   Y7263
• • • • • • • • • • • • • • I2a2a1b2a2a3a1a   Y7265
• • • • • • • • • • • • • I2a2a1b2a2a3a2   Y5692
• • • • • • • • • • • • I2a2a1b2a2a3b   Y10659
• • • • • • • • • • I2a2a1b2a2b   S25383, L1425
• • • • • • • • • • • I2a2a1b2a2b1   CTS5332.2/PF7472.2
• • • • • • • • • • • I2a2a1b2a2b2   Y4769
• • • • • • • • • • I2a2a1b2a2c   Y7426
• • • • • • •  • I2a2a1b2a3   L1290, L1317
• • • • • • •  • I2a2a1b2a4   Y7202, Y7844
• • • • • • • • I2a2a1b2b   L623, L147.3
• • • • • • • • I2a2a1b2c   CTS11871/S3673
• • • • • • • • • I2a2a1b2c1   Y10671
• • • • • I2a2a2   S9403/SK1254, Y6098
• • • • • • I2a2a2a   L1228
• • • • • • I2a2a2b   Y6099
• • • • I2a2b   L38/S154, L39/S155, L40/S156, L65.1/S159.1, L272.3
• • • • • I2a2b1   L533/S295
• • • • • I2a2b2   S2606
• • • • • • I2a2b2a   S24121
• • • • • • • I2a2b2a1   Y16415
• • • • • • • • I2a2b2a1a   Y16417
• • • • • • • • I2a2b2a1b   Y17117
• • • • • • I2a2b2b   FGC29612^^/Y13070^^
• • • • • • • I2a2b2b1   S2488, Y13074
• • • • • • • I2a2b2b2   PH1237
• • • • • • • I2a2b2b3   Y18919
• • • • • • • • I2a2b2b3a   Y18921
• • • • • • • • • I2a2b2b3a1   Y19084
• • • • • • • • • • I2a2b2b3a1a   Y19211
• • • • • • • • • • I2a2b2b3a1b   Y20202
• • I2bL415/S435, L416, L417/S332
• • I2c   L596/PF6907/S292, L597/S333, FGC18085/Y5340, FGC18140/Y5336, M3934/S6610/PF3934, PF3889, PF3893, PF3895, PF3896, PF3897, PF3898/S6659, PF3903/S3330, PF3911/S6728, PF3922, PF3929/S6694, PF3931, S6601, S6631/SK1266, S11759, S12355, Y5332, Y5338
• • I2c~ BY19, BY20, BY21, BY22/FGC18100, BY24, BY25, BY26, BY27, BY29, PF3881/S6673
• • • I2c1   PF3892, PF3904, PF3918, PF3924, PF3933
• • • • I2c1a   BY433
• • • • • I2c1a1   L1251.1, BY2776, BY2777, S9234, S10141, S11571, S16199, S16446, S17221, S17911, S18042, S18299, S21750, S22170, S22470, S23078, S26491
• • • • • • I2c1a1a   FGC18564, S14736
• • • • • • • I2c1a1a1   BY2789, BY2790, BY2791, BY2792
• • • • • I2c1a2   S6687, S6741
• • • • • • I2c1a2a   CTS4092
• • • • • • • I2c1a2a1   CTS7767.1
• • • • • • • • I2c1a2a1a   BY23/S6625
• • • • • • • • • I2c1a2a1a1   PF6328
• • • • • • • • • • I2c1a2a1a1a   F2044
• • • I2c2   BY133.1^/Y5127.1^, PH3539, Y16659, Y16660, Y16668, Y16672, Y16673, Y16675, Y16677, Y16678, Y16680, Y16682, Z26376, Z26380, Z26383, Z26384^, Z26386, Z26388, Z26389, Z26390, Z26396^, Z26399, Z26401, Z26403^, Z26404
• • • I2c2~Y16427
• • • • I2c2a   Z26381
• • • • • I2c2a1   Y16666
• • • • • • I2c2a1a   FGC6702.2/Y14376.2
• • • • • • I2c2a1b~Y16423

Notes

• Identical SNPs that were discovered separately are listed in alphabetical order, not necessarily in the order of discovery, and separated by "/". Examples: M307.1/P203.1, M450/S109, M438/P215, P41.2/M359.2, M436/P214/S33, P222/U250, L126/S165, L137/S166, L38/S154, L39/S155, L40/S156, and L65/S159.
• Page76: published accounts place this downstream from M170, which appears to be incorrect. A haplogroup J man has tested Page76- as expected, but three I1 males were Page76- as well. No known I male has been tested for Page76. Therefore Page76 is possibly in I2, but definitely not at the I node.
• Page123: The precise location of Page123 is uncertain. It is derived in four men with Haplogroup I1 SNPS and ancestral for one man in Haplogroup J. It has not been tested in Haplogroup I2.
• L69/S163 - Removed from I in 2011 and IJK in 2012, frequently mutating.
• Mutations previously on tree but found so far only in one man or in closely related men:
• S79: One confirmed results downstream from P30; considered a private snp, was previously listed as I1e. Listed 26 Sep 2008.
• L592: Out of 15 men tested, only three, all with the Straub surname, were derived. This snp is downstream from Z140/Z141. Listed February 15, 2012.
• L211: Downstream from L64 et al. As of 16 April 2012, 219 men have tested from L211, of which only two of the same surname have been confirmed as L211+
• M21:One confirmed tester in Underhill et al (197), downstream for L64 et al. As of 17 April 2012, 969 men in the FTDNA I1 Project have tested and all are ancestral.
• M507/P259: One confirmed result in Karafet et al (2008) downstream from L64 35 al. As of 17 April 2012, 679 men in the FTDNA I1 Project have tested and all are ancestral.
• M379: Downstream of M223, only one instance of M379+ is known, a Pakistani male in Polarity and Temporality of High-Resolution Y-ChromosomeDistributions in India Identify Both Indigenous and Exogenous Expansions and Reveal Minor Genetic Influence of Central Asian Pastoralists by Sengupta et al. The Pakistani sample was derived for M223. In the FTDNA I2b1/M223 Project, 237 men have tested for M379 and all were ancestral. Listed 28 June 2012.
• M161/PF4447 is downstream of L160. Listed 5 August 2012.
• PF6479 is approximately PF3827. Listed 18 December 2014, but PF3827 does not exist on tree.

Y-DNA haplogroup I is a European haplogroup, representing nearly one-fifth of the population. It is almost non-existent outside of Europe, suggesting that it arose in Europe. Estimates of the age of haplogroup I suggest that it arose prior to the last Glacial Maximum.
The two main subgroups of haplogroup I likely divided approximately 28,000 years ago:

• I1-M253 et al has highest frequency in Scandinavia, Iceland, and northwest Europe. In Britain, haplogroup I1-M253 et al is often used as a marker for "invaders," Viking or Anglo-Saxon. The I1b-M227 subclade is concentrated in eastern Europe and the Balkans and appears to have arisen in the last one thousand to five thousand years. It has been reported in Germany, Czech Republic, Poland, Estonia, Ukraine, Switzerland, Slovenia, Bosnia, Macedonia, Croatia, and Lebanon.
• I2-M438 et al includes I2* which shows some membership from Armenia, Georgia and Turkey; I2a1-P37.2, which is the most common form in the Balkans and Sardinia. I2a1a-M26 is especially prevalent in Sardinia. I2a2-M436 et al reaches its highest frequency along the northwest coast of continental Europe. I2a2a-M223 et al occurs in Britain and northwest continental Europe. I2a2a1-M284 occurs almost exclusively in Britain, so it apparently originated there and has probably been present for thousands of years.

References:
Athey et al, Resolving the Placement of Haplogroup I-M223 in the Y-chromosome Phylogenetic Tree. (pdf) Journal of Genetic Genealogy, 1:54-55, 2005.
Battaglia et al, Y-chromosomal Evidence of the Cultural Diffusion of Agriculture in Southeast Europe European Journal of Human Genetics, 249. 2008.
Behar et al, Genome-Wide Structure of the Jewish People. Nature, 446:238-42, 2010.
Bosch et al, Paternal and Maternal Lineages in the Balkans Show a Homogeneous Landscape over Linguistis Barriers except for the Isolated Aromuns. Annals of Human Genetics, 70:459-87, (2006).
Cinnioglu et al, Excavating Y-chromosome Haplotype Strata in Anatolia. (pdf) Human Genetics. 114:127-148, 2004.
El Sibai et al, Geographical Structure of the Y-Chromosomal Genetic Landscape of the Levant: A Coastal Inland Contrast, Annals of Human Genetics, 73:568-81, 2009. (abstract)
Francalacci et al, Low-Pass DNA Sequencing of 1200 Sardinians Reconstructs European Y-Chromosome Phylogeny. Science: Vol. 341 no. 6145, pp. 565-569, DOI: 10.1126/science.1237947, 2 August 2013.
Ghiani, et al, Population data for Y-chromosome haplotypes defined by AmpFlSTR YFiler PCR amplification kit in North Sardinia (Italy), Collegium Antropologicum, 33 (2): 643-51, 2009.
Karafet et al, New Binary Polymorphisms Reshape and Increase Resolution of the Human Y-Chromosomal Haplogroup Tree. Abstract. Genome Research, published online April 2, 2008. Supplementary Material.
Rootsi et al, Phylogeography of Y-Chromosome Haplogroup I Reveals Distinct Domains of Prehistoric Gene Flow In Europe. American Journal of Human Genetics, 75:128-137, 2004.
Rozen et al, Remarkably Little Variation in Proteins Encoded by the Y Chromosome's Single-Copy Genes, Implying Effective Purifying Selection. American Journal of Human Genetics, 85:923-928, 2009.
Scheinfeldt et al, Unexpected NRY Chromosome Variation in Northern Island Melanesia. Society for Molecular Biology, 2006.
Sims et al, Sub-Populations Within the Major European and African Derived Haplogroups R1b3 and E3a Are Differentiated by Previously Phylogenetically Undefined Y-SNPs. Human Mutation: Mutation in Brief #940, Online, 2007.
Underhill et al, New Phylogenetic Relationships for Y-chromosome Haplogroup I: Reappraising its Phylogeography and Prehistory in Rethinking the Human Evolution, Mellars P, Boyle K, Bar-Yosef O, Stringer C, Eds. McDonald Institute for Archaeological Research, Cambridge, UK, pp. 33-42, 2007b. (not available online)
Underhill et al, The Phylogeography of Y Chromosome Binary Haplotypes and the Origins of Modern Human Populations. (pdf) Annals of Human Genetics, 65:43-62, 2001.
Underhill et al, Y Chromosome Sequence Variation and the History of Human Populations. (pdf) Nature Genetics, 26: 358-361, 2000.
Y Chromosome Consortium (YCC), A Nomenclature System for the Tree of Human Y-Chromosomal Binary Haplogroups. Genome Research, 12:339-348, 2002.
Additional Resources: ISOGG Wiki - What you need to know about Genetic Genealogy. Founder Haplotypes (Modal Haplotypes for Y-Haplogroup I Varieties), Ken Nordtvedt. Deep Ancestry of I1a DYS19=16 Cluster, Phil Goff. I1b* Project Database, Timothy Weakley. Family Tree DNA Projects: Haplogroup I Y-DNA Project, Bob Waterous, Bernie Cullen, Paul Stone. I1 Y-DNA Haplogroup Project, Phil Goff, Aaron J. Hill, Steve Transgrud, Yme Drost, Ken Nordtvedt, William Hartley and Paul Stone. I1>Z58+ and I1>Z63+ Project, Aaron Salles Torres, Paul Stone, William Hartley, Wim Penninx. I1d - L22/S142 Project, Yme Drost, Albert Hulse. I1d2 (L205+) Project, Yme Drost. The I1-L1302 Project, Peter Sjölund, Jakob Norstedt-Moberg I1-Z140 YDNA Project, William Hartley, Wim Penninx and Aleksandr Shtrunov. Y-Haplogroup I2a Project, Bernie Cullen, Bill Morrow, Ken Nordtvedt, Zdenko Markovic. I2b2 L38+ Project, Timothy Weakley, Hans De Beule, Ken Nordtvedt. I2b L415 I2c L596 Haplogroup Project, Russ Livingstone, Bob May, Peter Hrechdakian, Ken Nordtvedt, Hovann Simonian. Iberian I1 Project, Aaron Salles Torres. I-L161 (I2a-Isles), Robert Gabel. I-M223 Y-Haplogroup Project, Aaron Salles Torres, Wayne Roberts, Ken Nordtvedt, George Anthony, Scott Engel Stewart. The I1d1 (I-P109) yDNA Haplogroup Project, Rebekah Canada, Nicolas Taban. The I1-L300 and Subclades yDNA Haplogroup Project, Artem Chuprov. The I-L1248 and Subclades yDNA Haplogroup Project, Jakob Norstedt-Moberg. I1 Eastern Europe, Aleksandr Shtrunov.
Corrections/Additions made since 1 January 2016:

• Moved BY211, BY212, BY213, BY214, BY215, BY216, BY217, BY218, BY220, BY222, BY223, BY224, BY225, BY226, BY228, BY230, BY232, BY233, BY234, BY235, BY236, BY240, BY244, BY245, BY269, BY270, BY271, BY272, BY273, BY274, BY275, BY276, BY277, BY278, BY19, BY20, BY21, BY22/FGC18100, BY24, BY25, BY26, BY27, BY29, BY148, BY18, BY128, BY12, BY13, A815, A813, A814, A811, A493, A376, M359.2/P41.2, PF3881/S6673, L1450, L1451, L1452, Z2037, Z2038, Z2040, L96/S196, L1224, CTS7362/S4713/Z2539, L1274, L1275, Z2041, Z119, Z125, S294/Z106, Z109, Z112, Z124, L673, L387.2, L700, L705.1/S465.1, L706.1, S336/Z75, S297/Z718, L234, L410, L242 from SNPs under Investigation to Tree Investigation; moved PF5479 from SNPs under Investigation to Notes; removed L96/S196 has nonsensical information on 7 January 2016.
• Added BY133.1/Y5127.1, PH3539, Y16659, Y16660, Y16668, Y16672, Y16673, Y16675, Y16677, Y16678, Y16680, Y16682, Z26376, Z26380, Z26383, Z26384, Z26386, Z26388, Z26389, Z26390, Z26396, Z26399, Z26401, Z26403, Z26404 on 13 January 2016.
• Added PF3664/Z2638, Z26381, Y16666, Y14976.2/FGC6702.2 to tree on 14 January 2016.
• Added Y16423, Y16427 to tree investigation on 17 January 2016.
• Added V2110.1 to tree on 14 February 2016.
• Removed CTS3529/PF3720/Z2649, CTS4039/PF3723/Z2650, CTS8959/Z2659, CTS9380/PF3784, PF3801/Z2663, PF3802/Z2664 from tree due to erroneous entries; removed BY417 from tree not meeting quality guidelines on 17 February 2016; moved L415/S435, L416, L417/S332 from tree to tree investigation.
• Removed S465.1 wrong mutation information for haplogroup I on 20 Mar 2016.
• Added Y16415, Y16417, Y17117 to tree on 5 April 2016.
• Added FGC29612/Y13070, PH1237, Y18919, Y18921, Y19084, Y19211, Y20202 to tree on 12 April 2016.
• Added AM00084/CTS2044, CTS10/Z2676, CTS22/Z2677, CTS40/Z2678, CTS136/Z2679, CTS209/V1092/Z2680, CTS565/Z2700, CTS571/Z2701, CTS641/Z2702, CTS733/Z2703, CTS765/Z2704, CTS883/Z2705, CTS1220/Z2706, CTS1266/Z2707, CTS1393/Z2708, CTS1739/Z2748, CTS1748/Z2749, CTS1755/Z2750, CTS1805/Z2752, CTS2375/Z2754, CTS2410/Z2755, CTS2644/Z2757, CTS2732/Z2758, CTS2738/Z2759, CTS2970/Z2760, CTS3268/Z2762, CTS3474/V2712/Z2764, CTS3506/Z2765, CTS3762/Z2766, CTS3843/Z2767, CTS4037/V2902/Z2768, CTS4130/V2932/Z2769, CTS4189/Z2770, CTS4295/Z2771, CTS4327/Z2772, CTS4342/Z2773, CTS4344/Z2774, CTS4387, CTS4400/Z2775, CTS4508/Z2776, CTS4532/Z2777, CTS4612/Z2778 to tree on 25 April 2016.
• Added CTS4656/Z2779, CTS4675/Z2780, CTS4790/Z2781, CTS5167/Z2784, CTS5247/Z2785, CTS5408/Z2786, CTS5513/Z2787, CTS5643/Z2790, CTS5705/Z2791, CTS5891/Z2793, CTS5993/Z2794, CTS6006/Z2795, CTS6022/Z2796, CTS6076/Z2797, CTS6109/Z2798, CTS6140/Z2799, CTS6221/Z2802, CTS6395/Z2803, CTS6547/Z2804, CTS6629/Z2805, CTS6765/Z2806, CTS6992, CTS7061/Z2807, CTS7121/Z2808, CTS7267/Z2809, CTS7386/Z2810, CTS7477/Z2811, CTS7652/Z2812, CTS7751/Z2813, CTS7949/Z2814, CTS8170/Z2816, CTS8224/Z2817, CTS8394/Z2818, CTS8652/Z2820, CTS8708/Z2821, CTS8716/Z2822, CTS9010/Z2824, CTS9258/Z2825, CTS9288/Z2826, CTS9368/Z2827, CTS9405/Z2828, CTS9486/Z2829, CTS9487/Z2830, CTS9518/Z2831, CTS9648/Z2832, CTS9749/Z2833, CTS9900, CTS10062/Z2834, CTS10140/Z2837, CTS10338, CTS10613/Z2840, CTS11036/Z2863 to tree on 26 April 2016.
• Added CTS11091/Z2865, CTS11126/Z2866, CTS11166/Z2867, CTS11214/Z2868, CTS11503/Z2869, CTS11526/Z2870, CTS11534/Z2871, CTS11552/Z2872, CTS11751/Z2874, CTS11775/Z2875, CTS11783/Z2876, CTS11838/Z2879, CTS11950/Z1880, CTS12008/Z2881, F3033.2/M1443.2/V4056.2/Z2838.2, FGC2422/Z2715, FGC2425/Z2714, FGC2427/Z2713, FGC2430/Z2699, FGC2431/Z2710, FGC2432/V1339/Z2712, FGC2435/Z2698, FGC2436/Y1879, FGC2439/Z2711, FGC2440/V1427/Y3658, FGC2471/Y1882, FGC2483/Y1933, L495/Z2788 to tree on 27 April 2016.
• Added L509/Z2864, L575/V4059/Z2839, L740/V3959/YSC0000296/Z2836, L750/YSC0000292/Z2725, L764/YSC0000271/Z2849, L841/YSC0000257/Z2751, L842/YSC0000264/Z2789, L843/YSC0000273/Z2878, L848/YSC0000299/Z2877, Page123/Z2763, S22865/Z2845, V1423/Z2717, V1771/Z2724, V2264/Z2728, Y1886, YSC0000259/Z2761, YSC0000301/Z2882, Z2718, Z2719, Z2721, Z2722, Z2723, Z2726, Z2727, Z2729, Z2730, Z2731, Z2733, Z2736, Z2737, Z2738, Z2740, Z2741, Z2742, Z2743, Z2744, Z2747, Z2823, Z2841, Z2842, Z2843, Z2844, Z2846, Z2847, Z2848, Z2850, Z2851, Z2852, Z2853, Z2854, Z2855, Z2856, Z2857, Z2861, Z2862, Z2885, Z2887 to tree on 28 April 2016.
• Added ZS2286 to tree; removed M1460/PF3816 not meeting quality guidelines on 30 April 2016.
• Added V2740 to tree on 9 May 2016.
• Added L300 and L1248 as projects on 28 June 2016.
• Added Y18119/Z17925, S2304/Z17937, Z17943, Z41181, Z41182, Z41183, Z41184, Z41185 on 11 July 2016.

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